  {"id":4163,"date":"2012-12-05T00:00:14","date_gmt":"2012-12-05T06:00:14","guid":{"rendered":"https:\/\/uwm.edu\/field-station\/?p=4163"},"modified":"2017-05-15T19:53:05","modified_gmt":"2017-05-16T00:53:05","slug":"daddy-longlegs-revisited","status":"publish","type":"post","link":"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/daddy-longlegs-revisited\/","title":{"rendered":"Daddy Longlegs Revisited (Family Phalangiidae)"},"content":{"rendered":"<p>Howdy, BugFans,<\/p>\n<p>The BugLady is busy and offers this rerun from 4 \u00bd years ago. The original has been enhanced with a few new, educational tidbits.<\/p>\n<h3>Daddy Longlegs<\/h3>\n<p>Daddy longlegs, a.k.a. Harvestman, (because the first species to be described were collected during harvest time, and because they are numerous then) are in the phylum Arthropoda (jointed legs), which consists of the classes Crustacea, Insecta and Arachnida and which, according to one author, accounts for about 80% of \u201cliving creatures.\u201d Don\u2019t let the eight legs and the spider-like appearance fool you; Daddy longlegs are not true spiders. Under the big umbrella of the class Arachnida, they belong in the order Opiliones, not with the true spiders in the order Araneae (there are some nice Greek myths about Ariadne and Arachne, and the BugLady trusts that BugFans will look them up on their own).<\/p>\n<p><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs09-6brz.jpg\" alt=\"d-longlegs09-6brz\" width=\"449\" height=\"414\" class=\"aligncenter size-full wp-image-4164\" srcset=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs09-6brz.jpg 449w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs09-6brz-300x277.jpg 300w\" sizes=\"auto, (max-width: 449px) 100vw, 449px\" \/><\/p>\n<p>DLLs have had the same basic design for the past 400 million years, give or take. They are made up of two body segments, a <em>cephalothorax<\/em> (fused head and body) and an abdomen. The DLL\u2019s waist is broad, leading one author to say it resembles a rice krispies on legs (they can move pretty fast on those long legs). They hear via vibrations; their eyes are on short stalks that look like antennae (but arachnids don\u2019t have antennae); and their eyesight is poor. The senses of taste, touch and smell are incorporated in the longer, second pair of their four pairs of legs. A startled DLL may wave these sensory legs in the air. When they clean themselves, which is often, they pay special attention to the second pair of legs. Males are smaller-bodied and brighter-colored. They are found in fields and meadows, both nocturnally and on bright days.<\/p>\n<p><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs-17rz.jpg\" alt=\"d-longlegs-17rz\" width=\"718\" height=\"441\" class=\"aligncenter size-full wp-image-4165\" srcset=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs-17rz.jpg 718w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs-17rz-300x184.jpg 300w\" sizes=\"auto, (max-width: 718px) 100vw, 718px\" \/><\/p>\n<p>A DLL might shed a leg if a predator grabbed it or if a venomous invertebrate stung\/bit it (it\u2019s called \u201cleg autotomy\u201d). They\u2019ve even developed a unique muscle attachment that minimizes bleeding when they do have to jettison an appendage. One researcher reporting in the <em>Journal of Arachnology<\/em> noted that the resulting \u201closs of fitness\u201d is still preferable to \u201ca catastrophic loss of fitness (e.g. death).\u201d DLLs cannot regenerate missing legs, and if they lose both of the sensory legs, they\u2019re toast. Despite their long legs, they carry their bodies close to the ground.<\/p>\n<p><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs-group-rz.jpg\" alt=\"d-longlegs-group-rz\" width=\"798\" height=\"388\" class=\"aligncenter size-full wp-image-4166\" srcset=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs-group-rz.jpg 798w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs-group-rz-300x146.jpg 300w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/d-longlegs-group-rz-768x373.jpg 768w\" sizes=\"auto, (max-width: 798px) 100vw, 798px\" \/><\/p>\n<p>Eisner, in <em>Secret Weapons<\/em>, calls their ability to produce a noxious odor their second line of defense. When they\u2019re grabbed by the body, a nasty-smelling chemical oozes out of a few dorsal glands, coats the body, and repels predators, especially ants. The odor is undetectable to humans, except those who really get up close and personal. <em>Phalangiists<\/em> claim they can distinguish species by their odor (Phalangiists. Scientists who work with daddy longlegs. In the Family Phalangiidae. OK, The BugLady may have made up that word).<\/p>\n<p>While not what you\u2019d call \u201csocial,\u201d DLLs do gather in large numbers&mdash;of hundreds and even thousands&mdash;for a variety of reasons; when they collect in fall, it\u2019s thought to be a warming strategy. A congregation of DLLs is a thicket of tiny knees.<\/p>\n<p><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/dddy-longlegs-mites12-2rz.jpg\" alt=\"dddy-longlegs-mites12-2rz\" width=\"820\" height=\"421\" class=\"aligncenter size-full wp-image-4167\" srcset=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/dddy-longlegs-mites12-2rz.jpg 820w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/dddy-longlegs-mites12-2rz-300x154.jpg 300w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/dddy-longlegs-mites12-2rz-768x394.jpg 768w\" sizes=\"auto, (max-width: 820px) 100vw, 820px\" \/><\/p>\n<p>Females lay as many as 40 pale green eggs underground in fall, using a long ovipositor. Head-of-a-pin-sized young hatch in spring, live one summer, mate, lay eggs and die. They produce neither web nor nest.<\/p>\n<p><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/dddy-lng-lgs-mate-rz.jpg\" alt=\"dddy-lng-lgs-mate-rz\" width=\"739\" height=\"597\" class=\"aligncenter size-full wp-image-4168\" srcset=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/dddy-lng-lgs-mate-rz.jpg 739w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/dddy-lng-lgs-mate-rz-300x242.jpg 300w\" sizes=\"auto, (max-width: 739px) 100vw, 739px\" \/><\/p>\n<p>Harvestmen are predators, eating insect eggs, small insects like aphids and springtails and critters as large as snails, earthworms, and other DLLs. A few species scavenge dead or decaying matter. Lacking the venomous fangs of true spiders, the Harvestman hunts for soft-bodied prey which it squeezes with its pincers and then stuffs into its mouth. Please note, boys and girls \u2013 \u201cLacking the venomous fangs of true spiders\u201d means that Daddy Longlegs &#8220;Do Not Bite.&#8221; They do not have fanged mouthparts and They! Do! Not! Bite!!<\/p>\n<p>The BugLady sometimes sees a daddy longlegs with its body resting flat on a leaf and its legs dangling over the sides, but she has no clue why it does that. She also has a (bad) slide of a shed skin of a DLL (shedding is another time in its life when a DLL might break off a leg)&mdash;the critter simply pulled itself out of a slit on the back and walked away, leaving empty legs and an empty body, intact.<\/p>\n<p>The BugLady is not sure why the pair of daddy longlegs on the window screen was \u201clip-locked\u201d one night in late summer, though in some species, the male gifts the female with a \u201csecretion\u201d before mating. Whatever was going on, there were nine voyeuristic DLLs nearby, watching.<\/p>\n<p>Relatively little is known about Harvestmen. Why? Because although they are beneficial in gardens, they are neither crop pests nor disease vectors, so when research funds are handed out, Phalangiists get bumped to the end of the line.<\/p>\n<p>Jonathan Swift said \u201ca flea hath smaller fleas that on him prey,\u201d (the curse of a liberal arts education). The tiny, red mites found on daddy longlegs fill the same niche as a mosquito or tick does on larger organisms. The Bug Lady always wonders what she would see if she could photograph the mite\u2019s leg.<\/p>\n<h3>Long-Bodied Cellar Spiders<\/h3>\n<p>Long-Bodies Cellar Spiders (<em>Pholcus phalangioides<\/em>, family Pholcidae) are true spiders and common household residents. Like the rest of us, they lived in caves before civilization and buildings arrived, and like the rest of us, the relative warmth provided by houses has allowed them to extend their range north. LBCSs are often mistaken for DLLs because of their long legs and similar size&mdash;in fact, a common name of these cellar spiders is \u201cDaddy Long-legs Spiders\u201d (another name, because of the shape of the cephalothorax, is Skull Spider).<\/p>\n<p><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/long-bodied-cellar-spider-3.jpg\" alt=\"long-bodied-cellar-spider-3\" width=\"1042\" height=\"685\" class=\"aligncenter size-full wp-image-4169\" srcset=\"https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/long-bodied-cellar-spider-3.jpg 1042w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/long-bodied-cellar-spider-3-300x197.jpg 300w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/long-bodied-cellar-spider-3-768x505.jpg 768w, https:\/\/uwm.edu\/field-station\/wp-content\/uploads\/sites\/380\/2016\/12\/long-bodied-cellar-spider-3-1024x673.jpg 1024w\" sizes=\"auto, (max-width: 1042px) 100vw, 1042px\" \/><\/p>\n<p>Unlike DLLs, LBCSs do spin webs, for the purpose of hunting (and they hang upside down in their webs). This spider spun its randomly-shaped web in the BugLady\u2019s shower; most of its confreres do live in her basement, but the BugLady\u2019s house wears Halloween decorations year \u2018round. When alarmed, LBCSs may shake or pump up and down so rapidly that both web and spider become a blur (some DLLs also bob, minus the web), confusing predators (but not, alas, the BugLady and her vacuum cleaner).<\/p>\n<p>Rule of Thumb: If it\u2019s outside, it\u2019s a DLL; if it\u2019s inside, it\u2019s a LBCS (especially now that the BugLady\u2019s had her back door replaced).<\/p>\n<p>Small World Department&mdash;Wikipedia shows a picture of a DLL eating the tail of a blue-tailed skink. Skinks shed their tails when grabbed just like DLLs shed legs.<\/p>\n<p>Folklore&mdash;Harvestmen point in the direction of cows. Around here, they\u2019re right.<\/p>\n<p>&nbsp;<br \/>\n<em>The Bug Lady<\/em><\/p>\n","protected":false},"excerpt":{"rendered":"<p><strong>Daddy Longlegs<\/strong> or Harvestman are predators, eating insect eggs, small insects like aphids and springtails and critters as large as snails, earthworms, and other DLLs. A few species scavenge dead or decaying matter. Lacking the venomous fangs of true spiders, the Harvestman hunts for soft-bodied prey which it squeezes with its pincers and then stuffs into its mouth. <\/p>\n","protected":false},"author":845,"featured_media":0,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":{"_acf_changed":false,"footnotes":"","uwm_wg_additional_authors":[]},"categories":[8],"tags":[310],"class_list":["post-4163","post","type-post","status-publish","format-standard","hentry","category-bug-of-the-week","tag-arachnids"],"yoast_head":"<!-- This site is optimized with the Yoast SEO Premium plugin v27.3 (Yoast SEO v27.3) - https:\/\/yoast.com\/product\/yoast-seo-premium-wordpress\/ -->\n<title>Field Station<\/title>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/uwm.edu\/field-station\/bug-of-the-week\/daddy-longlegs-revisited\/\" \/>\n<meta property=\"og:locale\" content=\"en_US\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Daddy Longlegs Revisited (Family Phalangiidae)\" \/>\n<meta property=\"og:description\" content=\"Daddy Longlegs or Harvestman are predators, eating insect eggs, small insects like aphids and springtails and critters as large as snails, earthworms, and other DLLs. 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